Magnolia has received keen interests from many botanists because it is believed one of early-diverging angiosperms in the evolutionary history of angiosperms. Although recent molecular phylogenetic studies showed that Magnolia is placed at the “Magnoliids” clade, a sister to eudicots + Ceratophyllales/Chloranthales (below tree), clarification of the feature of genome in Magnolia will provide a key to understand the evolutionary gap between basal angiosperms and eudicots.